Jonathan L Atwood (PhD)

Adjunct Faculty
Department of Environmental Studies

Selected Publications


Akçakaya, H. R., and J. L. Atwood. 1997. A habitat-based metapopulation model of the California Gnatcatcher. Conservation Biology 11:422-434.

Atwood, J. L. 1980c. The United States distribution of the California Black-tailed Gnatcatcher. Western Birds 11: 65-78.

Atwood, J. L. 1988. Speciation and geographic variation in black-tailed gnatcatchers. Ornithol. Monogr. No. 42.

Atwood, J. L. 1991. Subspecies limits and geographic patterns of morphological variation in California Gnatcatchers (Polioptila californica). Bull. S. Calif. Acad. Sci. 90: 118-133.

Atwood, J.L. 1992b. A maximum estimate of the California Gnatcatcher's population size in the United States. Western Birds 23: 1-10.

Atwood, J. L. 1992c. A closer look: California gnatcatcher. Birding 24:228-234.

Atwood, J.L. 1993a. California Gnatcatchers and coastal sage scrub: the biological basis for endangered species listing. Pp. 149-169, in J. E. Keeley [ed.], Interface Between Ecology and Land Development in California. S. Calif. Acad. Sci.; Los Angeles, California.

Atwood, J.L. 1993b. The politics of protection. Living Bird 12: 14-21.

Atwood, J.L. 1994. A Century of Avifaunal Change in Western North America: Endangered small landbirds of the western United States. Studies in Avian Biology 15: 328-339.

Atwood, J.L. and D.R. Bontrager. 2001. California Gnatcatcher (Polioptila californica). Pp. 1-32, in A Poole and F. Gill [eds.], The Birds of North America, No. 574. The Birds of North America, Inc.; Philadelphia, PA.

Atwood, J.L., and J.S. Bolsinger. 1992. Elevational distribution of California Gnatcatchers in the United States. J. Field Ornithol. 63: 159-168.

Atwood, J.L., and R.F. Noss. 1994. Gnatcatchers and development: a "train wreck" avoided? illahee 10: 123-130.

Zink, R.F., G.F. Barrowclough, J.L. Atwood, and R.C. Blackwell-Rago. Genetics, taxonomy, and conservation of the threatened California gnatcatcher. Conservation Biology 14: 1394-1405.


Akçakaya, H. R., J.L. Atwood, D. Breininger, C. T. Collins and B. Duncan. 2003. Metapopulation dynamics of the California Least Tern. J. Wildlife Management 67: 829-842.

Atwood, J. L. 1986. Delayed nocturnal occupation of breeding colonies by Least Terns (Sterna antillarum). Auk 103: 242-244.

Atwood, J. L., and P. R. Kelly. 1984. Fish dropped on breeding colonies as indicators of Least Tern food habits. Wilson Bull. 96: 34-47.

Atwood, J. L., and B.W. Massey. 1988. Site fidelity of Least Terns in California. Condor 90: 389-394.

Atwood, J. L., and D.E. Minsky. 1983. Least Tern foraging ecology at three major California breeding colonies. Western Birds 14: 57-72.

Massey, B.W., and J.L. Atwood. 1978. Plumages of the Least Tern. Bird-Banding 49: 360-371.

Massey, B.W., and J.L. Atwood. 1981. Second-wave nesting of the California Least Tern: age composition and reproductive success. Auk 98: 596-605.

Massey, B.W., D.W. Bradley, and J.L. Atwood. 1992. Demography of a California Least Tern colony including effects of the 1982 - 1983 El Niño. Condor 94: 976-983.

Thompson, B.C., J.A. Jackson, J. Burger, L.A. Hill, E.M. Kirsch, and J.L. Atwood. 1997. Least Tern (Sterna antillarum). In The Birds of North America, No. 290 (A. Poole and F. Gill, eds.). The Academy of Natural Sciences, Philadelphia, PA, and The American Ornithologists' Union, Washington, D.C.


Atwood, J.L. 1992a. Inferred destinations of spring migrant Common Yellowthroats based on plumage and morphology. Pp. 377-383, in J.M. Hagan and D.W. Johnston [eds.], Ecology and Conservation of Neotropical Migrant Landbirds. Smithsonian Institution Press, Washington.

Hagan, J.M., T.L. Lloyd-Evans, and J.L. Atwood. 1991. The relationship between latitude and the timing of spring migration of North American landbirds. Ornis Scandinavica 22: 129-136.

Hagan, J.M., T.L. Lloyd-Evans, J.L. Atwood, and D.S. Wood. 1992. Long-term changes in migratory landbirds in the northeastern United States: evidence from migration capture data. Pp. 115-130, in J.M. Hagan and D.W. Johnston [eds.], Ecology and Conservation of Neotropical Migrant Landbirds. Smithsonian Institution Press, Washington.

Lloyd-Evans, T. L. and J.L. Atwood. 2004. 32 years of changes in passerine numbers during spring and fall migrations in coastal Massachusetts. Wilson Bull. 116: 1-16.


Atwood, J.L., C. C. Rimmer, K.P. McFarland, S.H. Tsai, and L.R. Nagy. 1996. Distribution of Bicknell's Thrush in New England and New York. Wilson Bull. 108: 650-661.

Rimmer, C.C., J. L. Atwood, K.P. McFarland, and L.R. Nagy. 1996. Population density, vocal behavior, and recommended survey methods for Bicknell's Thrush. Wilson Bull. 108: 639-649.

Rimmer, C.C., J.L. Atwood, and L.R. Nagy. 1993. Bicknell's Thrush: a northeastern songbird in trouble? Bird Observer 21: 84-89.


Atwood, J.L. and C.T. Collins. The Island Scrub-Jay: origins, behavior, and ecology. Birding 29: 476-485.

Atwood, J. L., M. J. Elpers and C. T. Collins. 1990. Survival of breeders in Santa Cruz Island and mainland Scrub Jay populations. Condor 92: 783-788.

Atwood, J. L. 1980a. Breeding biology of the Santa Cruz Island Scrub Jay. Pp. 675-688, in D. M. Power [ed.], The California Islands: proceedings of a multidisciplinary symposium. Santa Barbara Natural History Museum, Santa Barbara, California.

Atwood, J. L. 1980b. Social interactions in the Santa Cruz Island Scrub Jay. Condor 82: 440-448.


Atwood, J.L., V.L. Fitz, and J.E. Bamesberger. 1991. Temporal patterns of activity at leks of the White-bellied Emerald. Wilson Bull. 103: 373-386.